Frederic Clements
Frederic Edward Clements (
1874-
1945) was an
American plant ecologist and pioneer in the study of
vegetation succession.
Born in
Lincoln, Nebraska, he studied
botany at the
University of Nebraska,
graduating in
1894 and obtaining a
doctorate in
1898. (One of his teachers was botanist
Charles Bessey, and he was a classmate of
Willa Cather and
Roscoe Pound.) While at the University of Nebraska, he met his future wife,
Edith Gertrude Schwartz (1874-1971), also a botanist and ecologist.
In
1907 he was appointed professor for
plant physiology at the University of Nebraska. From 1907 he was Professor of
botany at the
University of Minnesota in
Minneapolis. From 1917 to 1941 he was employed as an ecologist at the
Carnegie Institution of Washington in
Washington, D.C., where he was able to carry out dedicated ecological research.
During
winter he worked at
research stations in
Tucson,
Arizona and
Santa Barbara, California, while in the
summer he performed
fieldwork at the Carnegie's "
Alpine Laboratory," a research station in
Angel Canyon on the slopes of
Pikes Peak,
Colorado. During this time he worked alongside staff of the
U.S. Soil Conservation Service. In addition to his
field investigations, he carried out
experimental work in the
laboratory and
greenhouse, both at the Pikes Peak station and at Santa Barbara.
From his observations of the vegetation of Nebraska and the western United States, Clements developed one of the most influential theories of vegetation development. Vegetation cover does not represent a permanent condition but gradually changes over time. Clements suggested that the development of vegetation can be understood as a sequence of stages resembling the development of an individual organism. After a complete or partial disturbance, vegetation grows back (under ideal conditions) towards a mature "
climax state," which describes the vegetation best suited to the local conditions. Though any actual instance of vegetation might follow the ideal sequence towards climax, it can be interpreted in relation to that sequence, as a deviation from it due to non-ideal conditions.
Clements's climax theory of vegetation dominated plant ecology during the first decades of the twentieth century, though it was criticized significantly by ecologists
Henry Gleason and
Arthur Tansley early on, and by
Robert Whittaker mid-century, and largely fell out of favor. However, significant Clementsian trends in ecology re-emerged towards the end of the twentieth century.
In addition to botany and ecology, his research interests covered the
systematics of
fungi.
Among his works are:
*
The Phytogeography of Nebraska (1898; second edition, 1900)
*
Research Methods in Ecology (1905)
*
Plant Physiology and Ecology (1907)
*
Plant Succession (1916)
*
Plant Succession and Indicators (1928, reprinted 1973)
*
Flower Families and Ancestors (1928, with
Edith Clements)
*
Plant Ecology (1929, with J.E. Weaver)
*
The Genera of Fungi (1931, repr. 1965, with C. L. Shear)
In 1903, the flower
Clementsia rhodantha ("Clements' rose flower"), a
stonecrop, was named in honor of Frederic and Edith Clements. Clements died in Santa Barbara in 1945.